Image by Artem Maltsev

HÄSTARS NATURLIGA BETEENDEBEHOV – REFERENSER

För oss är det viktigt att certifieringen för Etisk hästhållning utgår ifrån relevant forskning om hästar. Det ska också finnas transparens kring vilken forskning som används som underlag till certifieringen. Här nedan följer därför en tematisk referenslista med alla de forskningsartiklar som ligger till grund för vårt arbete.


Känner du till forskning som saknas från denna lista? Tipsa oss gärna om lämpliga forskningsartiklar genom att mejla till oss på etiskhasthallning@gmail.com. Tack!


Grovfoder


Baumgartner, M., Boisson, T., Erhard, M. H., & Zeitler-Feicht, M. H. (2020). Common Feeding Practices Pose A Risk to the Welfare of Horses When Kept on Non-Edible Bedding. Animals, 10(3), 411.


Benhajali, H., Richard-Yris, M.-A., Ezzaouia, M., Charfi, F., & Hausberger, M. (2009). Foraging opportunity: a crucial criterion for horse welfare? Animal, 3(9), 1308-1312.


Boyd, L., & Bandi, N. (2002). Reintroduction of takhi, Equus ferus przewalskii, to Hustai National Park, Mongolia: time budget and synchrony of activity pre- and post-release. Applied Animal Behaviour Science, 78(2), 87-102.


Burla, J.-B., Ostertag, A., Patt, A., Bachmann, I., & Hillmann, E. (2016). Effects of feeding management and group composition on agonistic behaviour of group-housed horses. Applied Animal Behaviour Science, 176, 32-42.


Carslake, H. B., Pinchbeck, G. L., & McGowan, C. M. (2021). Equine metabolic syndrome in UK native ponies and cobs is highly prevalent with modifiable risk factors. Equine Veterinary Journal, 53(5), 923-934.


Correa, M. G., Rodrigues e Silva, C. F., Dias, L. A., da Silva Rocha Junior, S., Thomes, F. R., Alberto do Lago, L., de Mattos Carvalho, A., & Faleiros, R. R. (2020). Welfare benefits after the implementation of slow-feeder hay bags for stabled horses. Journal of Veterinary Behavior, 38, 61-66.


Davis, K. M., Iwaniuk, M. E., Dennis, R. L., Harris, P. A., & Burk, A. O. (2020). Effects of grazing muzzles on behavior and physiological stress of individually houses grazing miniature horses. Applied Animal Behaviour Science, 231.


Dierendonck, M. C. van, Bandi, N., Batdorj, D., Duegerlham, S., & Munkhtsog, B. (1996). Behavioural observations of reintroduced Takhi or Przewalski horses (Equus ferus przewalskii) in Mongolia. Applied Animal Behaviour Science, 50(2), 95-114.


Dosi, M. C. M., Kirton, R., Hallsworth, S., Keen, J. A., & Morgan, R. A. (2020). Inducing weight loss in native ponies: is straw a viable alternative to hay? The Veterinary Record, 187(8), e60.


Ellis, A. D., Fell, M., Luck, K., Gill, L., Owen, H., Briars, H., Barfoot, C., & Harris, P. (2015). Effect of forage presentation on feed intake behaviour in stabled horses. Applied Animal Behaviour Science, 165, 88-94.


Ellis, A. D., Redgate, S., Zinchenko, S., Owen, H., Barfoot, C., & Harris, P. (2015). The effect of presenting forage in multi-layered haynets and at multiple sites on night time budgets of stabled horses. Applied Animal Behaviour Science, 171, 108-116.


Fleurance, G., Duncan, P., & Mallevaud, B. (2001). Daily intake and the selection of feeding sites by horses in heterogeneous wet grasslands. Animal Research, 50(2), 149-156.


Frank, N., Geor, R. J., Bailey, S. R., Durham, A. E., & Johnson, P. J. (2010). Equine Metabolic Syndrome. Journal of Veterinary Internal Medicine, 24(3), 467–475.


Furugren, B., Ternrud, I., North, M., & Rundgren, M. (2013). Hästen. Natur & Kultur.


Glunk, E. C., Hathaway, M. R., Weber, W. J., Sheaffer, C. C., & Martinson, K. L. (2014). The Effect of Hay Net Design on Rate of Forage Consumption When Feeding Adult Horses. Journal of Equine Veterinary Science, 34(8), 986-991.


Jansson, A., Harris, P., Larsdotter Davey, S., Luthersson, N., Ragnarsson, S., & Ringmark, S. (2021). Straw as an Alternative to Grass Forage in Horses – Effects on Post-Prandial Metabolic Profile, Energy Intake, Behaviour and Gastric Ulceration. Animals, 11(2197), 2197.


Jansson, A., Gunnarsson, V. Þ., Ringmark, S., Ragnarsson, S., Söderroos, D., Ásgeirsson, E., Jóhannsdóttir, T. R., Liedberg, C., & Stefánsdóttir, G. J. (2021). Increased body fat content in horses alters metabolic and physiological exercise response, decreases performance, and increases locomotion asymmetry. Physiological Reports, 9(11).


Jonsson, H., & Egenvall, A. (2006). Prevalence of gastric ulceration in Swedish Standardbreds in race training. Equine Veterinary Journal, 38(3), 209-213.


King, S. R. (2002). Home range and habitat use of free-ranging Przewalskii horses at Hustai National Park, Mongolia. Applied Animal Behaviour Science, 78(2), 103-113.


Luthersson, N., Hou Nielsen, K., Harris, P., & Parkin, T. D. H. (2009). Risk factors associated with equine gastric ulceration syndrome (EGUS) in 201 horses in Denmark. Equine Veterinary Journal, 41(7), 625-630.


Maisonpierre, I. N., Sutton, M. A., Harris, P., Menzies-Gow, N., Weller, R., & Pfau, T. (2019). Accelerometer activity tracking in horses and the effect of pasture management on time budget. Equine Veterinary Journal, 51(6), 840-845.


Mayes, E., & Duncan, P. (1986). Temporal patterns of feeding behaviour in free-ranging horses. Behaviour, 96(1/2), 105-129.


Raspa, F., Roggero, A., Palestrini, C., Marten Canavesio, M., Bergero, D., & Valle, E. (2021). Studying the Shape Variations of the Back, the Neck, and the Mandibular Angle of Horses Depending on Specific Feeding Postures Using Geometric Morphometrics. Animals, 11(763), 763.


Rochais, C., Henry, S., & Hausberger, M. (2018). “Hay-bags” and “Slow feeders”: Testing their impact on horse behaviour and welfare. Applied Animal Behaviour Science, 198, 52-59.


Sigurjonsdottir, H., Thorhallsdottir, A. G., Hafthorsdottir, H. M., & Granquist, S. M. (2012). The Behaviour of Stallions in a Semiferal Herd in Iceland: Time Budgets, Home Ranges, and Interactions. International Journal of Zoology, 2012.


Sykes, B. W., Hewetson, M., Hepburn, R. J., Luthersson, N., & Tamzali, Y. (2015). European College of Equine Internal Medicine Consensus Statement – Equine Gastric Ulcer Syndrome in Adult Horses. Journal of Veterinary Internal Medicine, 29(5), 1288-1299.


Thorne, J. B., Goodwin, D., Kennedy, M. J., Davidson, H. P. B., & Harris, P. (2005). Foraging enrichment for individually housed horses: Practicality and effects on behaviour. Applied Behaviour Science, 94(1), 149-164.


Wichert, B., Nater, S., Wittenbrink, M. M., Wolf, P., Meyer, K., & Wanner, M. (2008). Judgement of hygienic quality of roughage in horse stables in Switzerland. Journal of Animal Physiology and Animal Nutrition, 92(4), 432-437.



Sociala behov


Berger, J. (1977). Organizational systems and dominance in feral horses in the Grand Canyon. Behavioral Ecology and Sociology, 2(2), 131.


Bourjade, M., des Roches, A. de B., & Hausberger, M. (2009). Adult-Young Ratio, a Major Factor Regulating Social Behaviour of Young: A Horse Study. PLoS ONE, 4(3), 1-5.


Bourjade, M., Moulinot, M., Henry, S., Richard-Yris, M.-A., & Hausberger, M. (2008). Could adults be used to improve social skills of young horses, Equus caballus? Developmental Psychobiology, 50(4), 408-417.


Christensen, J. W., Ladewig, J., Søndergaard, E., & Malmkvist, J. (2002). Effects of individual versus group stabling on social behaviour in domestic stallions. Applied Animal Behaviour Science, 75(3), 233-248.


Christensen, J. W., Søndergaard, E., Thodberg, K., & Halekoh, U. (2011). Effects of repeated regrouping on horse behaviour and injuries. Applied Animal Behaviour Science, 133(3), 199-206.


Flauger, B., & Krueger, K. (2013). Aggression level and enclosure size in horses (Equus caballus). Pferdeheilkunde, 29(4), 495-504.


Fureix, C., Bourjade, M., Henry, S., Sankey, C., & Hausberger, M. (2012). Exploring aggression regulation in managed groups of horses Equus caballus. Applied Animal Behaviour Science, 138(3-4), 216-228.


Grogan, E. H., & McDonnell, S. M. (2005). Injuries and blemishes in a semi-feral herd of ponies. Journal of Equine Veterinary Science, 25(1), 26-30.


Heitor, F., do Mar Oom, M., & Vicente, L. (2006). Social relationships in a herd of Sorraia horses: Part II. Factors affecting affiliative relationships and sexual behaviours. Behavioural Processes, 73(3), 231-239.


Jørgensen, G. H. M., Borsheim, L., Meidell, C. M., Søndergaard, E., & Bøe, K. E. (2009). Grouping horses according to gender – Effects on aggression, spacing and injuries. Applied Animal Behaviour Science, 120(1), 94-99.


Kaseda, Y. (1981). The Structure of the Groups of Misaki Horses in Toi Cape. 日本畜産学会報, 52(3), 227.


Kaseda, Y., & Khalil, A. M. (1996). Harem size reproductive success of stallions in Misaki feral horses. Applied Animal Behaviour Science, 47(3), 163-173.


Keeling, L. J., Bøe, K. E., Christensen, J. W., Hyyppä, S., Jansson, H., Jørgensen, G. H. M., Ladewig, J., Meidell, C. M., Särkijärvi, S., Søndergaard, E., & Hartmann, E. (2016). Injury incidence, reactivity and ease of handling of horses kept in groups: A matched case control study in four Nordic countries. Applied Animal Behaviour Science, 185, 59-65.


Kerekes, V., Sándor, I., Nagy, D., Ozogány, K., Göczi, L., Ibler, B., Széles, L., & Barta, Z. (2021). Trends in demography, genetics, and social structure of Przewalski’s horses in the Hortobagy National Park, Hungary over the last 22 years. Global Ecology and Conservation, 25.


Kimura, R. (1998). Mutual grooming and preferred associate relationships in a band of free-ranging horses. Applied Animal Behaviour Science (Netherlands), 59(4), 265-276.


Knubben, J. M., Fürst, A., Gygax, L., & Stauffacher, M. (2008). Bite and kick injuries in horses: Prevalence, risk factors and prevention. Equine Veterinary Journal, 40(3), 219-223.


Lee, J., Floyd, T., Erb, H., & Houpt, K. (2011). Preference and demand for exercise in stabled horses. Applied Animal Behaviour Science, 130(3), 91-100.


Löckener, S., Reese, S., Erhard, M., & Wöhr, A.-C. (2016). Pasturing in herds after housing in horseboxes induces a positive cognitive bias in horses. Journal of Veterinary Behavior: Clinical Applications and Research, 11, 50-55.


Mendonça, R. S., Pinto, P., Inoue, S., Ringhofer, M., Godinho, R., & Hirata, S. (2021). Social determinants of affiliation and cohesion in a population of feral horses. Applied Animal Behaviour Science, 245.


Pacheco, M. A., & Herrera, E. A. (1997). Social structure of feral horses in the Llanos of Venezuela. Journal of Mammalogy, 78(1), 15.


Pierard, M., McGreevy, P., & Geers, R. (2019). Effect of density and relative aggressiveness on agonistic and affiliative interactions in a newly formed group of horses. Journal of Veterinary Behavior, 29, 61-69.


Ruet, A., Lemarchand, J., Parias, C., Mach, N., Moisan, M.-P., Foury, A., Briant, C., & Lansade, L. (2019). Housing Horses in Individual Boxes Is a Challenge with Regard to Welfare. Animals, 9(9), 621.


Roberts Jr., J. M., & Browning, B. A. (1998). Proximity and threats in highland ponies. Social Networks, 20(3), 227-238.


Sigurjónsdóttir, H., & Haraldsson, H. (2019). Significance of Group Composition for the Welfare of Pastured Horses. Animals, 9(1), 14.


Sigurjónsdóttir, H., Van Dierendonck, M. C., Snorrason, S., & Thórhallsdóttir, A. G. (2003). Social relationships in a group of horses without a mature stallion. Behaviour, 140(6), 783-804.


Suagee-Bedore, J. K., Linden, D. R., & Bennett-Wimbush, K. (2021). Effect of Pen Size on Stress Responses of Stall-Housed Horses Receiving One Hour of Daily Turnout. Journal of Equine Veterinary Science, 98.


Søndergaard, E., Jensen, M. B., & Nicol, C. J. (2011). Motivation for social contact in horses measured by operant conditioning. Applied Animal Behaviour Science, 132(3), 131-137.


Søndergaard, E., & Ladewig, J. (2004). Group housing exerts a positive effect on the behaviour of young horses during training. Applied Animal Behaviour Science, 87(1), 105-118.


Van Dierendonck, M. C., de Vries, H., Schilder, M. B. H., Colenbrander, B., Þorhallsdóttir, A. G., & Sigurjónsdóttir, H. (2009). Interventions in social behaviour in a herd of mares and geldings. Applied Animal Behaviour Science, 116(1), 67-73.


Vervaecke, H., Stevens, J. M. G., Vandemoortele, H., Sigurjónsdóttir, H., & De Vries, H. (2007). Aggression and dominance in matched groups of subadult Icelandic horses (Equus caballus). Journal of Ethology, 25(3), 239.


Visser, E. K., Ellis, A. D., & Van Reenen, C. G. (2008). The effect of two different housing conditions on the welfare of young horses stabled for the first time. Applied Animal Behaviour Science, 114(3), 521-533.


Yarnell, K., Hall, C., Royle, C., & Walker, S. L. (2015). Domesticated horses differ in their behavioural and physiological responses to isolated and group housing. Physiology & Behavior, 143, 51-57.



Rörelsefrihet


Boyd, L., & Bandi, N. (2002). Reintroduction of takhi, Equus ferus przewalskii, to Hustai National Park, Mongolia: time budget and synchrony of activity pre- and post-release. Applied Animal Behaviour Science, 78(2), 87-102.


Chaplin, S. J., & Gretgrix, L. (2010). Effect of housing conditions on activity and lying behaviour of horses. Animal, 4(5), 792-795.


Chaya, L., Cowan, E., & McGuire, B. (2006). A note on the relationship between time spent in turnout and behaviour during turnout in horses (Equus caballus). Applied Animal Behaviour Science, 98(1), 155-160.


Hampson, B. A., de Laat, M. A., Monot, J., Bailliu, D., & Pollitt, C. C. (2013). Adaption of horses to a novel dynamic feeding system: movement and behavioural responses. Equine Veterinary Journal, 45(4), 481-484.


Hampson, B. A., Morton, J. M., Mills, P. C., Trotter, M. G., Lamb, D. W., & Pollitt, C. C. (2010). Monitoring distances travelled by horses using GPS tracking collars. Australian Veterinary Journal, 88(5), 176-181.


Hoffmann, G., Bentke, A., Rose-Meierhöfer, S., Berg, W., Mazetti, P., & Hardarson, G. H. (2012). Influence of an active stable system on the behavior and body condition of Icelandic horses. Animal, 6(10), 1684-1693.


Janczarek, I., Kedzierski, W., Stachurska, A., & Wilk, I. (2016). Can Releasing Racehorses to Paddocks be Beneficial? Heart Rate Analysis – Preliminary Study. Annals of Animal Science, 16(1), 87-97.


Jørgensen, G. H. M., & Bøe, K. E. (2007). A note on the effect of daily exercise and paddock size on the behaviour of domestic horses (Equus caballus). Applied Animal Behaviour Science, 107(1), 166-173.


Keleman, Z., Grimm, H., Vogl, C., Long, M., Cavalleri, J. M. V., Auer, U., & Jenner, F. (2021). Equine Activity Time Budgets: The Effect of Housing and Management Conditions on Geriatric Horses and Horses with Chronic Orthopaedic Disease. Animals, 11(1867), 1867.


King, S. R. (2002). Home range and habitat use of free-ranging Przewalskii horses at Hustai National Park, Mongolia. Applied Animal Behaviour Science, 78(2), 103-113.


Lee, J., Floyd, T., Erb, H., & Houpt, K. (2011). Preference and demand for exercise in stabled horses. Applied Animal Behaviour Science, 130(3), 91-100.


Lesimple, C., Reverchon-Billot, L., Galloux, P., Stomp, M., Boichot, L., Coste, C., Henry, S., & Hausberger, M. (2020). Free movement: A key for welfare improvement in sport horses? Applied Animal Behaviour Science, 225.


Maisonpierre, I. N., Sutton, M. A., Harris, P., Menzies-Gow, N., Weller, R., & Pfau, T. (2019). Accelerometer activity tracking in horses and the effect of pasture management on time budget. Equine Veterinary Journal, 51(6), 840-845.


Rivera, E., Benjamin, S., Nielsen, B., Shelle, J., & Zanella, A. J. (2002). Behavioral and physiological responses of horses to initial training: the comparison between pastured versus stalled horses. Applied Animal Behaviour Science, 78(2), 235-252.


Rose-Meierhöfer, S., Klaer, S., Ammon, C., Brunsch, R., & Hoffmann, G. (2010). Activity Behavior of Horses Housed in Different Open Barn Systems. Journal of Equine Veterinary Science, 30(11), 624-634.


Ruet, A., Arnould, C., Levray, J., Lemarchand, J., Mach, N., Moisan, M.-P., Foury, A., Briant, C., & Lansade, L. (2020). Effects of a temporary period on pasture on the welfare state of horses housed in individual boxes. Applied Animal Behaviour Science, 228.


Sigurjonsdottir, H., Thorhallsdottir, A. G., Hafthorsdottir, H. M., & Granquist, S. M. (2012). The Behaviour of Stallions in a Semiferal Herd in Iceland: Time Budgets, Home Ranges, and Interactions. International Journal of Zoology, 2012.


Stachurska, A., Różańska-Boczula, M., & Wnuk-Pawlak, E. (2021). The difference in the locomotor activity of horses during solitary and paired release. Pferdeheilkunde, 37(1), 50-54.


Van Dierendonck, M. C., Bandi, N., Batdorj, D., Duegerlham, S., & Munkhtsog, B. (1996). Behavioural observations of reintroduced Takhi or Przewalski horses (Equus ferus przewalskii) in Mongolia. Applied Animal Behaviour Science, 50(2), 95-114.


Werhahn, H., Hessel, E. F., Schulze, H., & Van den Weghe, H. F. A. (2011). Temporary Turnout for Free Exercise in Groups: Effects on the Behavior of Competition Horses Housed in Single Stalls. Journal of Equine Veterinary Science, 31(7), 417-425.


Werhahn, H., Hessel, E. F., & Van den Weghe, H. F. A. (2012). Competition Horses Housed in Single Stalls (I): Behaviour and Activity Patterns during Free Exercise According to Its Configuration. Journal of Equine Veterinary Science, 32(1), 45-52.


Werhahn, H., Hessel, E. F., & Van den Weghe, H. F. A. (2012). Competition Horses Housed in Single Stalls (II): Effects of Free Exercise on the Behavior in the Stable, the Behavior during Training, and the Degree of Stress. Journal of Equine Veterinary Science, 32(1), 22-31.



Lösdrift med ligghall


Anderson, U. S., Benne, M., Bloomsmith, M. A., & Maple, T. L. (2002). Retreat space and human visitor density moderate undesirable behavior in petting zoo animals. Journal of Applied Animal Welfare Science: JAAWS, 5(2), 125-137.


Hillyer, M. H., Taylor, F. G. R., Proudman, C. J., Edwards, G. B., Smith, J. E., & French, N. P. (2002). Case control study to identify risk factors for simple colonic obstruction and distension in horses. Equine Veterinary Journal, 34(5), 455-463.


Kaya, G., Sommerfeld-Stur, I., & Iben, C. (2009). Risk factors of colic in horses in Austria. Journal of Animal Physiology and Animal Nutrition, 93(3), 339-349.


Kjellberg, L., Yngvesson, J., Sassner, H., & Morgan, K. (2021). Horses’ Use of Lying Halls and Time Budget in Relation to Available Lying Area. Animals, 11(3214), 3214.


Mazzola, S. M., Colombani, C., Pizzamiglio, G., Cannas, S., Palestrini, C., Costa, E. D., Gazzonis, A. L., Bionda, A., & Crepaldi, P. (2021). Do You Think I Am Living Well? A Four-Season Hair Cortisol Analysis on Leisure Horses in Different Housing and Management Conditions. Animals, 11(2141), 2141.


Owen, M. A., Swaisgood, R. R., Czekala, N. M., & Lindburg, D. G. (2005). Enclosure choice and well-being in giant pandas: is it all about control? Zoo Biology, 24(5), 475-481.


Reeves, M. J., Salman, M. D., & Smith, G. (1996). Risk factors for equine acute abdominal disease (colic): results from a multi-center case-control study. Preventive Veterinary Medicine, 26(3-4), 285-301.


Ross, S. R. (2006). Issues choice and control in the behaviour of a pair of captive polar bears (Ursus maritimus). Behavioural Processes, 73(1), 117-120.


Sauveroche, M., Henriksson, J., Theodorsson, E., Svensson Holm, A.-C., & Roth, L. S. V. (2020). Hair cortisol in horses (Equus caballus) in relation to management regimes, personality, and breed. Journal of Veterinary Behavior. 37, 1-7.


Yngvesson, J., Rey Torres, J. C., Lindholm, J., Pättiniemi, A., Andersson, P., & Sassner, H. (2019). Health and Body Conditions of Riding School Horses Housed in Groups or Kept in Conventional Tie-Stall/Box Housing. Animals, 9(3), 1-13.